diff --git a/bfd/elflink.c b/bfd/elflink.c
index c9d5da2dab4..6fe90412b28 100644
--- a/bfd/elflink.c
+++ b/bfd/elflink.c
@@ -1272,14 +1272,25 @@ _bfd_elf_merge_symbol (bfd *abfd,
       olddyn = (oldsec->symbol->flags & BSF_DYNAMIC) != 0;
     }
 
-  /* Handle a case where plugin_notice won't be called and thus won't
-     set the non_ir_ref flags on the first pass over symbols.  */
   if (oldbfd != NULL
-      && (oldbfd->flags & BFD_PLUGIN) != (abfd->flags & BFD_PLUGIN)
-      && newdyn != olddyn)
+      && (oldbfd->flags & BFD_PLUGIN) != (abfd->flags & BFD_PLUGIN))
     {
-      h->root.non_ir_ref_dynamic = true;
-      hi->root.non_ir_ref_dynamic = true;
+      if (newdyn != olddyn)
+	{
+	  /* Handle a case where plugin_notice won't be called and thus
+	     won't set the non_ir_ref flags on the first pass over
+	     symbols.  */
+	  h->root.non_ir_ref_dynamic = true;
+	  hi->root.non_ir_ref_dynamic = true;
+	}
+
+      if ((oldbfd->flags & BFD_PLUGIN) != 0
+	  && hi->root.type == bfd_link_hash_indirect)
+	{
+	  /* Change indirect symbol from IR to undefined.  */
+	  hi->root.type = bfd_link_hash_undefined;
+	  hi->root.u.undef.abfd = oldbfd;
+	}
     }
 
   /* NEWDEF and OLDDEF indicate whether the new or old symbol,
diff --git a/ld/testsuite/ld-plugin/lto.exp b/ld/testsuite/ld-plugin/lto.exp
index 68d2db1caeb..c343132c460 100644
--- a/ld/testsuite/ld-plugin/lto.exp
+++ b/ld/testsuite/ld-plugin/lto.exp
@@ -238,6 +238,15 @@ if { [at_least_gcc_version 10 0] } {
 	 "-flto -fno-common $lto_no_fat" \
 	 {pr25355.c} \
 	 [list [list "nm" "$plug_opt" "pr25355.d"]]] \
+	[list "pr28264.so" \
+	 "-shared -Wl,--version-script=pr28264.ver" \
+	 "-flto $lto_no_fat -fPIC" \
+	 {pr28264.c} \
+	 {{readelf {--dyn-syms --wide} pr28264-1.d} \
+	  {readelf {--dyn-syms --wide} pr28264-2.d} \
+	  {readelf {--dyn-syms --wide} pr28264-3.d} \
+	  {readelf {--dyn-syms --wide} pr28264-4.d}} \
+	 {pr28264.so}] \
     ]]
 }
 
diff --git a/ld/testsuite/ld-plugin/pr28264-1.d b/ld/testsuite/ld-plugin/pr28264-1.d
new file mode 100644
index 00000000000..5ff89694c98
--- /dev/null
+++ b/ld/testsuite/ld-plugin/pr28264-1.d
@@ -0,0 +1,5 @@
+Symbol table '\.dynsym' contains [0-9]+ entries:
+ +Num: +Value +Size Type +Bind +Vis +Ndx Name
+#...
+ +[0-9]+: +[0-9a-f]+ +[0-9a-f]+ +FUNC +GLOBAL +DEFAULT .*[0-9]+ _?foo@VERSION.1
+#pass
diff --git a/ld/testsuite/ld-plugin/pr28264-2.d b/ld/testsuite/ld-plugin/pr28264-2.d
new file mode 100644
index 00000000000..f2f629d4a32
--- /dev/null
+++ b/ld/testsuite/ld-plugin/pr28264-2.d
@@ -0,0 +1,5 @@
+Symbol table '\.dynsym' contains [0-9]+ entries:
+ +Num: +Value +Size Type +Bind +Vis +Ndx Name
+#...
+ +[0-9]+: +[0-9a-f]+ +[0-9a-f]+ +FUNC +GLOBAL +DEFAULT .*[0-9]+ _?bar@@VERSION.1
+#pass
diff --git a/ld/testsuite/ld-plugin/pr28264-3.d b/ld/testsuite/ld-plugin/pr28264-3.d
new file mode 100644
index 00000000000..b6a8f1f7848
--- /dev/null
+++ b/ld/testsuite/ld-plugin/pr28264-3.d
@@ -0,0 +1,4 @@
+#failif
+#...
+ +[0-9]+: +[0-9a-f]+ +[0-9a-f]+ +FUNC +GLOBAL +DEFAULT .*[0-9]+ _?foo
+#...
diff --git a/ld/testsuite/ld-plugin/pr28264-4.d b/ld/testsuite/ld-plugin/pr28264-4.d
new file mode 100644
index 00000000000..408b853831c
--- /dev/null
+++ b/ld/testsuite/ld-plugin/pr28264-4.d
@@ -0,0 +1,4 @@
+#failif
+#...
+ +[0-9]+: +[0-9a-f]+ +[0-9a-f]+ +FUNC +GLOBAL +DEFAULT .*[0-9]+ _?bar
+#...
diff --git a/ld/testsuite/ld-plugin/pr28264.c b/ld/testsuite/ld-plugin/pr28264.c
new file mode 100644
index 00000000000..4efa934fd3e
--- /dev/null
+++ b/ld/testsuite/ld-plugin/pr28264.c
@@ -0,0 +1,11 @@
+void
+__attribute__ ((symver ("foo@VERSION.1")))
+foo (void)
+{
+}
+
+void
+__attribute__ ((symver ("bar@@VERSION.1")))
+bar1 (void)
+{
+}
diff --git a/ld/testsuite/ld-plugin/pr28264.ver b/ld/testsuite/ld-plugin/pr28264.ver
new file mode 100644
index 00000000000..1608fcae92d
--- /dev/null
+++ b/ld/testsuite/ld-plugin/pr28264.ver
@@ -0,0 +1,8 @@
+VERSION.1
+{
+  global:
+    foo;
+    bar;
+  local:
+    *;
+};